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A Most Important Structure – The Skeleton (No. 3 Apr-97)

According to Darwinian evolution, the skeleton has evolved over millions of years to become efficient for delivering mechanical energy and for load support. The skeleton has many interesting design features: it is lightweight, it is able to sustain repetitive loads (fatigue resistance) and it can withstand impacts – most of the time. Some say it is an optimal structure, but optimal for what? For running around the forest? For driving a car?

Since the designer does not respond well to interrogation, we must try to ai???work backwardsai??i?? to find the answer. There are two significant constraints to be taken into account: (i) the skeleton must function throughout most of the construction phase (i.e. the growth phase); (ii) relatively weak and elastic bone must be used (Ultimate Stress = 100 N/mm2, Youngai??i??s modulus = 20 kN/mm2). Given these constraints, how is the efficiency achieved?

Firstly bones are self-repairing structures. Cells living inside the bone are continually resorbing it while others are forming it. Bones are alive! In this way, aged material is replaced by new “damage-free” tissue. This means that the actual strength of the tissue need not be as high as a fatigue analysis would suggest.

Secondly, the ai???cumulative memoryai??i?? contained in the DNA replicates the skeleton, but this replication process is partly in response to the mechanical environment. Mechanical loading is needed to run the correct genetic program.

In conclusion, perhaps a search for the “optimisation function” is looking for a rule which is not there. There is no such thing as an optimal bone, just one that balances, in some ai???fuzzyai??i?? way, the conflicts inherent in life itself.

Patrick J. Prendergast, Ph.D., C.Eng.
Trinity College, Dublin.

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